The TradeMark Security System (TMSS) was a feature of Sega Mega Drive motherboards.
Evidence suggests TMSS was originally intended as a region lockout system for the Mega Drive to keep Japanese games from running on foreign systems: the only games that intentionally omitted TMSS support were many early Japanese games (and two later ones); the few international games that omitted support did so due to development issues. Sega's official developer manual also refers to the system as the "US security code". However, this notion appears to have been dropped quickly, as early international models lack the feature and Japanese ROMs started shipping TMSS-compliant in mid-1989 (though the Sega Channel BIOS is an anomaly). It is unknown whether any Japanese Mega Drive models have TMSS.
In the US, it was eventually used to ensure games which were unlicensed by Sega (and thus not receiving the Sega Seal of Quality, which was attainable through a fee paid to Sega) were not playable. Though TMSS was not produced in whole until late 1992/early 1993, the necessary code was in programmer documentation as far back as 1989. Some early games that did not implement TMSS in full consequently did not work in newer board revisions.
TMSS works by simply requiring instances of the SEGA trademark to appear in various memory locations (see below). Should an unlicensed game appear on the market, Sega could exert its rights over the SEGA trademark and sue the developer for trademark infringement.
The legality of this system was challenged in the United States by Accolade in the court case SEGA vs. Accolade, which ruled in Accolade's favor. The verdict set a precedent that copyrights do not extend to non-expressive content in software that is required by another system to be present in order for that system to run the software. While the system was never challenged elsewhere, Sega did not enforce it after the Accolade case, and consequently several unlicensed game developers started popping up.
In order for software to run on a Mega Drive/Genesis with a TMSS built in, the software must do two things:
Absolute Entertainment, for some reason, faked the TMSS screen if it detected a non-TMSS board on all their Mega Drive games (including BattleTech, which was published under the Extreme Entertainment Group label which they had recently purchased from Micronet).